Belemnites
The cigar-shaped fossils of Belemnites are common in the West
Coast’s Jurassic and Cretaceous. The solid portion of the shell commonly found
represents a portion of the guard, devoid of the phragmocone and pro-ostracum.
The most massive element of a Belemnoid’s shell is the guard, or rostrum.
Unrepresented in the shell of any other cephalopod, the rostrum is fossilized as
calcite leading to durability and frequent preservation. The phragmocone is the
chambered portion of the shell and, as in
nautiloids and ammonites, there is a siphuncle. However; the belemnite siphuncle
is straight, without septal necks. Each septal chamber of a baculite, ammonite,
or nautiloid has shell encasing the siphuncle which is pinched as a collar or
neck at septal junctions. The pro-ostracum is a projection of the phragmocone’s
dorsal surface extending over the visceral body much like the pen in today’s
squid.
Belemnoids have been found from the Mississipian age, but true belemnites didn’t
evolve until the Triassic, and they persisted through to the Eocene, although
they are very rare after the Cretaceous. Exceptionally well preserved belemnites
from Europe have shown double rows of chitinoid hooks called onychites.
Onychites have been found in rows of eight and ten, generally interpreted as
arms with no long grasping tentacles. The internal structures of all belemnoids
are the same, but the animals’ proportions varied greatly. In the Triassic,
Ausseites had a small guard with a long phragmocone, while others like the
Jurassic’s Belemnoteuthis had a very thin guard with a short fat phragmocone.
Guards can be short or very long, or even diverting from the conical form. In
the Cretaceous

Hibolites, the guard is hastate, getting thin just before the phragmocone, while
Duvalia has a guard that flattens out like a vertical fin. Taxonomy in
belemnites is based on the shape of the rostrum, size, cross-section, and
grooves. Seen in cross-section the rostrum has rings that can be interpreted
much like those of a tree; the average belemnoid was thought to live about four
or five years. Based on the rostrum, the average belemnite animal was 300 to
500mm and no more than half a metre in length. Of course, there are also
extremes such as Neohibolites which was about 100 mm long, depending on its arm
length; and a rostrum from Indonesia is so big it suggests a belemnite that may
have been 5 meters long! Rarely found compared to the guard, the protoconch of
the average belemnite is a small ball at the phragmocone’s tip. Belemnoid
protoconchs from Harrison Lake are 3/4 of a millimeter across, and the
phragmocone has 40 septa. For comparison, a new-born ammonite from the same
locality is a millimeter in size with a protoconch of about the same size as the
belemnite. The small sized new-born belemnite would be reliant on the planktonic
world just as the ammonite, and the reduction of a large percentage of the
plankton at the end of the Cretaceous could have been responsible for at least
part of the
belemnites’ demise.The cigar-shaped fossils of Belemnites are common in the West
Coast’s Jurassic and Cretaceous. The solid portion of the shell commonly found
represents a portion of the guard, devoid of the phragmocone and pro-ostracum.
The most massive element of a Belemnoid’s shell is the guard, or rostrum.
Unrepresented in the shell of any other cephalopod, the rostrum is fossilized as
calcite leading to durability and frequent preservation. The phragmocone is the
chambered portion of the shell and, as in
nautiloids and ammonites, there is a siphuncle. However; the belemnite siphuncle
is straight, without septal necks. Each septal chamber of a baculite, ammonite,
or nautiloid has shell encasing the siphuncle which is pinched as a collar or
neck at septal junctions. The pro-ostracum is a projection of the phragmocone’s
dorsal surface extending over the visceral body much like the pen in today’s
squid.Belemnoids have been found from the Mississipian age, but true belemnites didn’t
evolve until the Triassic, and they persisted through to the Eocene, although
they are very rare after the Cretaceous. Exceptionally well preserved belemnites
from Europe have shown double rows of chitinoid hooks called onychites.
Onychites have been found in rows of eight and ten, generally interpreted as
arms with no long grasping tentacles. The internal structures of all belemnoids
are the same, but the animals’ proportions varied greatly. In the Triassic,
Ausseites had a small guard with a long phragmocone, while others like the
Jurassic’s Belemnoteuthis had a very thin guard with a short fat phragmocone.
Guards can be short or very long, or even diverting from the conical form.
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Figure 2. Cutaway of a belemnite showing the rostrum, phragmocone, and pro-ostracum in life position. |
In the Cretaceous Hibolites, the guard is hastate,
getting thin just before the phragmocone, while Duvalia has a guard that
flattens out like a vertical fin. Taxonomy in belemnites is based on the shape
of the rostrum, size, cross-section, and grooves. Seen in cross-section the
rostrum has rings that can be interpreted much like those of a tree; the average
belemnoid was thought to live about four or five years. Based on the rostrum,
the average belemnite animal was 300 to 500mm and no more than half a metre in
length. Of course, there are also extremes such as Neohibolites which was about
100 mm long, depending on its arm length; and a rostrum from Indonesia is so big
it suggests a belemnite that may have been 5 meters long! Rarely found compared
to the guard, the protoconch of the average belemnite is a small ball at the
phragmocone’s tip. Belemnoid protoconchs from Harrison Lake are 3/4 of a
millimeter across, and the phragmocone has 40 septa. For comparison, a new-born
ammonite from the same locality is a millimeter in size with a protoconch of
about the same size as the belemnite. The small sized new-born belemnite would
be reliant on the planktonic world just as the ammonite, and the reduction of a
large percentage of the plankton at the end of the Cretaceous could have been
responsible for at least part of the
belemnites’ demise.
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